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left many broken pieces of milfoil floating in the water col- <br />umn. The fact that there was a rapid decline in the biomass <br />in the 3rd week of sampling supports this interpretation. <br />Harvested areas had lower average biomass than refer- <br />ence areas for 6 weeks after harvest. Biomass increased in <br />harvested areas and decreased in reference areas such that <br />the two were no longer different 6 weeks after harvest based <br />on weekly comparisons of harvested and reference biomass <br />(Figure 2) [week 3 p = 0.0019, week 6 p = 0.0919, week 9 p= <br />0.31321. <br />Plant canopy cover was inversely related to plant biomass <br />(Figure 3), with plant biomass explaining 46% of the vari- <br />ability in the plant canopy ratios. As plants regrew after har- <br />vesting, plant canopy ratios showed a similar pattern to <br />biomass readings, with harvested and reference canopy <br />ratios converging to a single point at week 9 (Figure 4) . Har- <br />vested areas had significantly higher plant canopy ratios than <br />reference areas until the 6th week after harvesting [p=0.0022 <br />week 3, p=0.0820 week 6, p = 0.912 week 91. <br />Other researchers have found that harvesting reduced <br />biomass for only 3 to 4 weeks. Thus, milfoil populations har- <br />vested in mid-July attained pre -harvest and reference levels <br />within 23 days after harvesting in LaRue Reservoir, Ohio <br />(Cooke et al. 1990) and sites in Dundee and Herring Creek, <br />Maryland had to be harvested once per month throughout <br />the summer in order to attain effective management (Rawls <br />1975). However, harvesting is sometimes more effective. Mil - <br />foil in lake Wingra, Wisconsin never returned to reference <br />levels when harvested during July (Kimbel and Carpenter <br />1981). Similarly, in Lake Minnetonka, both plant canopy <br />ratio and biomass data indicated that harvesting was an effec- <br />tive control method for up to 6 weeks after harvesting, when <br />harvested in early July. <br />While harvested areas in Lake Minnetonka achieved ref- <br />erence area biomass and canopy density 6 weeks after harvest <br />they did not achieve a significantly higher biomass or canopy <br />density in the following weeks. Despite a higher relative <br />growth rate in harvested areas than in reference areas follow- <br />ing <br />ollowing harvest, harvested areas at no time attained higher biom- <br />ass or canopy density than reference areas (Figures 2 and 4). <br />The growth of milfoil is often limited by water depth (Aiken <br />et al., 1979) which could account for harvested areas' biom- <br />ass leveling out after regrowth. <br />Plant biomass in relation to environmental factors <br />The percentage of milfoil by weight in the reference <br />areas for each site was not correlated with any lake character- <br />istic, nor did the percentage of milfoil correlate with biomass <br />in the control areas (r2= 0.005). This indicates that in Lake <br />Minnetonka a high percentage of milfoil does not always <br />coincide with high total plant biomass nor does having high <br />total plant biomass imply that there is greater abundance of <br />milfoil in that area than in areas with lower plant biomass. <br />While milfoil relative abundance was not correlated with abi- <br />otic conditions, total plant biomass was correlated (p<0.01) <br />with water clarity and percent sediment organic matter <br />(Table 2). <br />There were significant differences in sediment organic <br />matter among sites (p<0.05) (Table 1). Within the range of <br />0.9 <br />• <br />r <br />0.8 <br />v <br />O S <br />•� U <br />N <br />a t 0.7 <br />o <br />c a <br />a m <br />U -a <br />c Y <br />o v <br />a. 0.6 <br />`c <br />0 <br />FL <br />v <br />R2 = 0.46 • • <br />• <br />0.5 <br />300 400 500 600 700 800 900 <br />Plant biomass (grams dry weight m-2) <br />Figure 3. Plant canopy ratio in relation to plant biomass. <br />physical variability in our plots, higher total plant biomass <br />occurred in sediment with more organic matter. This corre- <br />lation could be due to a preference of aquatic plants for sed- <br />iments which are enriched with organic matter or could be <br />caused, as Lillie and Barko (1990) suggest, by the enrich- <br />ment of the sediments by the aquatic vegetation itself. <br />WO <br />0.9 <br />r <br />a <br />0.8 <br />t <br />O U <br />U <br />O m <br />0.7 <br />O L <br />0 <br />c a <br />o a� <br />v a 0.6 <br />G y, <br />O <br />CL <br />0 0.5 <br />IL <br />0.4 <br />0.3 <br />• Unharvested <br />Harvested <br />Weeks after harvest <br />July August September <br />10 <br />Figure 4. Plant canopy ratio changes over time in harvested and unhar- <br />vested plots. Harvest done on July 3, 1990. Results presented as mean ± stan- <br />dard error. <br />J. Aquat. Plant Manage. 32: 1994. 59 <br />